Cole Elasser / USA TODAY Sports

Cal run defense returns to its old ways, allowing 6.5 yards per carry against Oregon State

Cal tallied the highest yards per carry rushing average of Sonny Dykes era in Saturday's overtime loss to Oregon State.

California running back Tre Watson rushed for a career-high 134 yards on 16 carries with two touchdowns (including his first rushing touchdown of the season). His previous career high was 110 rushing yards against Oregon State last year.

It was Watson's second career 100-yard rushing game. Bears running back Khalfani Muhammad, after missing last week's win over Utah, also finished with a career high rushing, totaling 165 yards on 21 carries, including a 50-yard touchdown rush in the fourth quarter. His previous best was 164 yards on the ground last year at Texas.

Cal's 317 rushing yards are a season high, topping the 189 the Bears had in their opener vs. Hawaii. The rushing total marks the highest in the Sonny Dykes era. The last time Cal ran for more yards was when the Bears had 318 rushing yards at Washington State in 2012.

Cal' 7.2 yards per rushing attempt mark the highest average yards per carry of the Dykes era.

The Bears' 44 rushing attempts tonight was the most since Cal ran 46 times against the Beavers on Nov. 14 of last year, when the Bears racked up 307 yards on the ground. It's tied for the fourth most rushing attempts (Portland State, 2013) in the Dykes era, with the 2014 game against the Beavers seeing the Bears run 47 times, and the 2013 game against Oregon seeing 51 Cal rushing attempts.

This was the second time this season (Texas - 307) that the Bears allowed more than 300 yards rushing. Cal allowed six drives of 70 yards or more against the Beavers. The Bears had allowed no more than four in any of the previous five games.

Cal and Oregon State combined to set a Cal record for most rushing yards by both teams in one game with 791 yards (Cal – 317, Oregon State – 474). The previous mark was 758 yards in a Cal-Pacific game ion Sept. 20, 1958 (Cal –350, Pacific – 408).

Over the last two weeks, the Bears had held opponents to 3.26 yards per rush. On Saturday, the Beavers averaged 6.5, the same as Cal allowed over the first three games of the season.

Oregon State quarterback Darell Garretson completed 13 of 24 passes for 85 yards and two interceptions, but rushed 10 times for 105 yards and two touchdowns.

Beavers running back Ryan Nall only played three quarters and change, but rushed for 221 yards on 14 carries, with three touchdowns.

Saturday's game was Cal's first to go into overtime since the Bears posted a 59-56 double OT victory over Colorado on Sept. 27, 2014 in Berkeley.

Linebacker Ray Davison -- who came into the game leading the Pac-12 in tackles (45) added seven more to his total. Davison returned his first career interception for a touchdown in the second quarter.

It was Cal's first interception returned for a touchdown since linebacker Devante Downs against Grambling State in 2015. Cameron Walker also returned an interception for a touchdown in that game against the Tigers. Cal's last defensive touchdown was a 47-yard fumble return by Stefan McClure against Washington State in 2015. Davison's 39-yard return is the longest by a Cal defender since Marc Anthony (61) vs. Southern Utah in 2012.

Quarterback Davis Webb rushed for his third touchdown of the season in the third quarter, which also gives him 25 total touchdowns in 2016. Webb injured his hand on the Bears' second possession, but played the rest of the game. No other Cal quarterback has thrown a single pass this season. Place kicker Matt Anderson made fields goals from 27, 29 and 39 yards, with the final one coming in over time. His first bounced off the cross bar after being tipped. Dating back to last season, he has made 25 of his last 27 field goal attempts.

Nickel back Trey Turner caught his second career interception in the second quarter.

Saturday was Cal's fifth straight game decided by 10 or fewer points.

Punter Dylan Klumph completed a pass to defensive end Evan Weaver on a fake punt in the first quarter. It was the first time a Cal punter completed a pass since Cole Leininger against Stanford in 2013.

Klumph finished the night with four punts for 159 yards, averaging 39.8 yards per punt, with a long of 57 and two inside the 20.

This was the first time since the first half against Stanford last year that the Cal offense did not score a touchdown in the first half.

Cal scored 20 consecutive points during one stretch – 17 in a row in the fourth quarter to send the game into overtime and the first three of the extra period.

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