In his fourth 100-yard game of the season – and ninth of his career – Wilson, a sophomore, had a career-high 11 catches for 126 yards.
Senior wide receiver Isiah Barton added another three catches for 18 yards.
As a result of such play, the Cougars' offense had 30 first downs, only the second time this season that it has had that many, and outgained Oregon's offense 462 yards to 454. The Ducks' offense entered the game sixth in the nation – and hadn't been outgained at home since barely escaping with a win over Purdue in 2009.
Granted, it could have been more. WSU had a number of drops. Other passes to open receivers were behind or low their intended target.
THE 24-YARD PASS from senior quarterback Marshall Lobbestael to Karstetter with 3:31 left in the third period cut the Cougars' deficit to 29-20 – and was the team's only touchdown reception of the game, which is hardly common in the age of passing versus running the ball.
But the Cougars were marred by repeated problems in the red zone, which, in turn, limited their scoring.
"You have to play almost perfect to beat a team like this," Karstetter said in a postgame radio interview. "You've got to execute in the red zone."
Lobbestael took the sole blame for his team's inability to do so.
"When I get down into the red zone, I've got to be smarter with the ball," he said in a postgame interview with the Eugene Register-Guard. "It's a team game, but at the same time the quarterback has total control over forcing it in there."
STILL, COACH Paul Wulff said he felt his team had a solid, balanced game plan.
"We had a few drops and misthrows that could've been big plays for us," he said in a postgame radio interview.
Karstetter took some responsibility on behalf of the receiving corps for some of their leader's inconsistencies. Lobbestael threw for 337 yards and a touchdown – and two interceptions, both when the Cougars were inside Oregon's 25-yard line. He was also sacked three times.
"It's a team thing when he gets sacked," he said. "We've got to get open."
Wulff was more succinct in his assessment of the offensive woes.
"We had so many opportunities," he said.
Despite the outcome, Karstetter seemed optimistic about some aspects of the Cougars' play.
"When you play a great team, you've got to make plays," he said. "I think we beat them in a lot of phases of the game today."
Horton is one away from tying the WSU record of five, set by Jason David in 2002.
"I saw one more person coming, and I felt my legs start giving out," Horton said of nearly being cut off by an Oregon offensive lineman near the Ducks' 25. "I would have been talked about all week."
"They did a good job of sorting stuff out – communicating in a loud, hostile environment," he said.